Thanks for sharing the export settings.
It seems as if the export issue is a bug related to Spine version 3.8.79 (incorrectly stripping some whitespace at a mesh), after upgrading to Spine 3.8.81, the problem no longer occurs.
Regarding your setup: Please note that you should not use meshes in form of a quad for your wings, you could as well use a normal Region Attachment, which would allow for automatic whitespace stripping. When you currently export your atlas textures, it contains lots of unnecessary whitespace due to using Mesh Attachments. Unless you are planning on editing the meshes later to fit the image, then of course ignore the lines above.